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Nicolas was born in Bronx and his dad was a respected Mafia member. At age eight, Aunt Mo told him to kill his parents and his younger sister and framing uncle Jimmy for the killings and kill him too. Mo adopted him and when turning into an adult she forced him to have sex with her, he didn't enjoy it because of his homosexuality and at this turned him a psychotic sadist and choked his aunt to death with a pillow during an intercourse. During sex she always shouted "Fuck Aunt Mo".
Later was employed by Don Cesare and was ordered to scare a Japanese man, David Kai, in order to join their Mafia. They dined and Kai didn't accept his offer, so Cavella revealed that the meat they eat was his two younger sons, then killed the elder two but left Kai alive. He expected it would make him popular but he was wrong. So he traveled to Boston with his best men, Pittsy, a psychotic henchman that got mad from some war and Ink a quiet guy who was called like that because he first killed with a pen. In Boston he ran drug business for fifteen years and started having relationships with corrupt military officer Rawlins.
Later Punisher almost exterminated the Mafia in New York, so Larry Barrucci called Cavella to help him restart the Mafia. Nicky agreed but first they had to get rid of Punisher, who at that time was being interrogated by Microchip who was working with the CIA to force him in working with the Government in counter-terrorism. Pittsy castrated Agent Roth to tell him where they kept Castle. He was kept in a hotel. They went in wounding and killing other agents including Micro. Pittsy was shot in his shoulder and Ink was killed by O'Brein. Micro and Frank escaped to one of his safehouses and Cavella sent an army of Mafiosos to get them and all failed. Pittsy too went to kill him but was impaled on a gate. So Cavella forced Barrucci to kill Punisher but was instantly shot in the thorax. Frank followed Nicky but was interrupted by a dying Pittsy. Pittsy was shot in the face. Later Cavella killed two cops to flee with their car.
After two years he was known as one of a few criminals to survive at confronting the Punisher. He became more dangerous when he and Teresa Gazzera, Pittsy's sister, profaned the Castle family graves and urinated on the bodies, this permitted him to be elected as the Don. Cavella stupidly thought that Punisher will head after him but instead he massacred local gangs. His former lover Bill Rawlins was assigned by CIA to stop Frank Castle, so he helped Cavella. The plan was that Rawlins would shoot Castle with a sniper while he's attacking Cavella who will be sitting on a chair as a bait. The plan fails because Rawlins' ex-wife, Kate alongside Agent Roth kidnap Rawlins and she starts to shoot at Cavella's men, saving Castle and taking him with them. Cavella and Teresa follow their car at Roth's apartment were Rawlins is interrogated. Teresa tries to seduce Nicky but he pushed her away, in order to like her she goes in but is killed by Castle and O'Brien who were having sexual intercourse. Rawlins kills Roth and escapes. The next day as Castle and O'Brien get out, they face Cavella who is holding a boy hostage. Castle warns him that if he hurts the boy he'll give him a suffering death, as soon as he hears this his cowardliness comes out and leaves the boy. He's put in the luggage booth and taken away in a forest were Frank shoots him in the belly while explaining him how he'll die. After three days he died a slow and painful death, probably because of blood poisoning. Hunters found him with his eyes pecked out by birds. He was forever remembered as Castle's most notable adversaries.