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Star Trek/Legion of Super-Heroes #5 finds one team of Starfleet officers and Legionnaires in the distant past, where they encounter the force that erased both their timelines from existence. Meanwhile, in an altered 23rd century, another team of Starfleet officers and Legionnaires comes face to face with the latter-day version of that same powerful force. Fully aware of how history was changed, both teams must confront the question of whether or not anything can be done to set things right.

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